braconid wasp

insect
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Also known as: Braconidae

braconid wasp, (family Braconidae), large and diverse family of more than 17,000 species of parasitic wasps, a number of which are useful for the biological control of insect pests. Like most species of solitary wasps, braconid wasps are parasitoids, meaning their larvae feed and develop on or within the bodies of other arthropods. They parasitize many insect groups, especially Lepidoptera (moths and butterflies), Coleoptera (beetles), Diptera (flies), and other Hymenoptera (antsbees, and wasps).

Taxonomy

See also list of ants, bees, and wasps.

Physical description

Braconid wasps are dark or dull in color and relatively small, seldom exceeding 1.5 cm (0.6 inch) in length. They are similar in appearance to ichneumon wasps, a related family of parasitic wasps, though braconids are typically smaller. The wings are sometimes banded or spotted. The ovipositor, or egg-laying organ, of the female is long and conspicuous; only a few species use the ovipositor as a stinger for defense.

As insects, braconid wasps have a body formed of a head, thorax, and abdomen, and six legs. Most species resemble other wasps in having a long, slender, curved abdomen with a fairly narrow waist. They have a pair of large compound eyes on the side of the head and three ocelli (simple eyes) on the top of the head. They usually differ from social wasps in having longer antennae with more segments, or sections.

Lion (panthera leo)
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Natural history and parasitism

Braconid larvae are either endoparasitic, living within their hosts, or ectoparasitic, living on their hosts. Endoparasitic adult females lay an egg or eggs in the larvae or eggs of the host (braconid wasps do not typically exploit adult hosts). Many species are egg-larval parasitoids, meaning the wasp eggs are laid within the host eggs but do not develop until the host has hatched. The braconid larva remains within the still-living host’s body at least until the host’s tissues and fluids are exhausted and the wasp is ready to pupate. The pupa may be formed in the body of the host, attached to the body of the host, or formed away from the host in a cocoon on a leaf or stem. In some species, as many as 150 larvae may develop in a single host. Certain groups of ectoparasitic females are idiobionts, paralyzing their hosts with their ovipositor before laying eggs on or near the host.

As adults, most braconid wasps feed on flower nectar and pollen and are at least incidental pollinators.

Major biological control species

Many species of braconids are valuable in the control of insect pests and are introduced and managed in many places as a form of biological control. Apanteles glomeratus, for example, parasitizes the larvae of the cabbage butterfly (Pieris rapae) and the cabbage looper (Trichoplusia ni), important pests of a number of Brassica crops. Apanteles congregatus parasitizes the tobacco hornworm (Manduca sexta) and the tomato hornworm (Manduca quinquemaculata).

Some braconids attack wood-boring pests such as beetles of the families Buprestidae and Cerambycidae. The braconid Chremilus rubiginosus attacks the grain weevil (Sitophilus granarius). In the Mediterranean region Opius concolor is a parasite of the olive fly (Dacus oleae), which is a destructive pest of commercial olives.

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The Editors of Encyclopaedia BritannicaThis article was most recently revised and updated by Melissa Petruzzello.